What is termite's biggest enemy?

Termite's biggest enemy is a combination of natural predators, environmental factors, and human pest control methods that target their need for moisture and cellulose, but in terms of biological enemies, ants are considered termites' biggest natural enemy. Ants and termites are natural competitors that often nest near each other and battle for territory, with many ant species including carpenter ants, fire ants, and army ants actively raiding termite colonies, killing workers, and stealing eggs and larvae, and some ant species have evolved specialized tactics to breach termite mounds. Other natural enemies include aardvarks, anteaters, and pangolins which dig into termite mounds and eat thousands of termites daily, plus birds, lizards, and spiders that prey on swarming termites during mating flights. Nematodes, which are microscopic parasitic worms, infect and kill termites from inside and are used as biological control. Fungi like Metarhizium anisopliae infect termites and spread through colonies. However, termites' biggest enemy overall in protecting homes is professional pest control using integrated methods, including liquid termiticide barriers containing fipronil that create a lethal zone in soil, bait systems with growth regulators that termites carry back to the colony, and moisture control because termites need moisture to survive, so fixing leaks, improving drainage, and removing wood-to-soil contact are critical enemies of termites. Homeowners are also enemies by reducing conducive conditions. So while ants are the biggest natural enemy, human prevention and treatment are the most effective enemies for protecting houses from over five billion dollars in damage annually in the US.